- 2003 PhD in Biomedical Sciences. Faculty of Veterinary Medicine, University of Zagreb. Thesis Title: Induction of Cytochrome P450 Enzymes of Fish by Oxytetracycline and Combination Ormetoprim-Sulfadimethoxine.
- 1997 MS in Oceanology. Faculty of Science, Graduate Study in Oceanology, University of Zagreb. Thesis Title: Bacteria Isolated from Fish Living in Vrana Lake from the Island of Cres.
- 1993 DVM (Doctor of Veterinary Medicine). Faculty of Veterinary Medicine, University of Zagreb. Thesis Title: Cross Antigens from Trichinella spiralis and Trichuris suis Nematode Protein Preparations.